In the phylogenetic trees, the United States and most South Korea haplotypes belonged to East Plain lineage, except one individual was classified into Loess Plateau lineage (Figure 2). All Japan haplotypes formed a monophyly group in Loess Plateau lineage, which was related to this single South Korea individual (Figure 2).
This shows that the origin of U.S. and Japan lanternflies both are from South Korea, which suggests another spread pattern: China --> South Korea --> Japan & U.S. This suggests that the spotted lanternflies in China only invaded South Korea, and South Korean spotted lanternflies were more invasive, targeting two other countries.